Mother Rachel Fee and her partner Nyomi have been found guilty of murdering Rachel's two-year-old son Liam in Fife.
The couple were convicted of assaulting and killing the young child at his home in Thornton in March 2014. They had blamed another boy for the murder.
They were also convicted of a catalogue of abuse against two other children.
The jury returned a majority verdict after a seven-week trial at the High Court in Livingston. The couple are due to be sentenced on 6 July.
The jury found that Rachel and Nyomi Fee had subjected the toddler to an escalating pattern of cruelty during his short life.
- The shocking abuse of Liam Fee
- Will lessons be learned from Liam's death?
- Horrific evidence endured by the jury
The couple, originally from Ryton, Tyne and Wear, remained impassive as the jury found them guilty of all the charges against them.
Liam Fee's father, Joseph Johnson, was in tears at the verdict and had to be consoled by friends.
The jury were excused from duty for 10 years.
In the days before his death, Liam suffered a broken leg and arm. He was killed by a blow to his abdomen which was so severe that it ruptured his heart.
In total, he was found to have more than 30 injuries.
The couple had blamed another boy for Liam's death. However, during several interviews with specially-trained officers and a social worker, it became clear that the boy had not strangled or suffocated Liam.
He had put his hand over the toddler's mouth several days before his death but Liam had been walking and talking afterwards.
Makeshift cage
The court heard that Nyomi had forced the same boy's fist into Liam's mouth after he died in order to leave DNA traces.
But the jury found Nyomi and Rachel to be jointly responsible for Liam's murder.
In addition, they subjected two other boys to a series of physical and psychological abuse over two years - including forcing one to sleep in a makeshift cage and another to sleep in a room with rats and snakes which they said "ate little boys".
Social services had been alerted to the Fees but had failed to take any action.
His nursery had alerted social services, worried by a change in Liam, and the fact that he was losing weight and had a number of injuries.
Liam's childminder had also made her concerns known.
A senior Fife social worker admitted to the court that at one point Liam "fell off their radar".
Fife Child Protection Committee is to carry out a significant case review.
Elaine Torrance, the president of Social Work Scotland, said such cases were "exceptional".
"Liam's death is an absolute tragedy and the level of cruelty that Liam experienced was deeply shocking," she said.
"What makes it worse is that the abuse was carried out by the very people Liam should have been able to trust the most: his mum and his step-mum."media playe
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